Except CPU speed and power, RAM quantity, disc read speed (likely), memory latency (it will trail there, DDR3 latency<GDDR5) etc...
I think, maybe someone more knowledgeable can explain better, but in a way the CPU will be the true limiting factor anyway. The GPU cannot actually create vertices/geometry or something like that. So it can improve some things yes, but not the most fundamental ones. Like a prettier coat of paint on the same building analogy.
So the PS4 alleged superior GPU can make some things look better, but not really fundamentally alter anything (if I'm correct)